V. N. Aliyassova, K. K. Akhmetov, I. R. Aspanova
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companions. Sabre-toothed cats, having underdeveloped molar teeth, could
not chew meat and thus could only eat the soft innards of their prey, so a
significant proportion of the meat they left behind went to the hyenas.
Fossilised hyenas found at Gussinyi Perelet were similar in type to modern
African spotted hyenas.
No less interesting a representative of this ancient fauna was the
rhinoceros, Chilotherium. There are currently two hypotheses regarding
their ecological niche. The first is based on the structure of the chewing
surfaces of their teeth, which is characteristic of animals grazing on the
tough grass of the steppe. Further support for this hypothesis is provided
by the middle toe of their feet, which suggests adaption to life in open
plains. The second hypothesis suggests that based on their short legs these
rhinoceroses dwelt in low wetland areas, and only later forms of this species
were able to adapt to the open steppe. It is believed that these rhinos lived
mainly in waterside areas and spent a major part of their existence close to
or in water, much like modern hippopotamuses.
The evolution of the landscape and geography of the Priirtyshja
region (in which Pavlovdar is located) had to be revaluated in 1958 after V.
V. Kuznetsov discovered a fragment of another continental plate typical of
the coastal area containing a rather large turtle carapace in the Gussinyi
Perelet sediments.
5
Further research of this coastal plate identified the
carapace as belonging to a type of sea turtle. This made a very interesting
find: Gussinyi Perelet is a long way inland, yet the remains of sea turtles
were discovered amongst the mammalian bones of the Hipparion fauna.
The discovery of marine fauna in a continental area is an unusual
phenomenon, requiring explanation. There is reason to believe that in the
era the Hipparion fauna roamed, the territory now known as Kazakhstan
had a climate similar to modern subtropical regions such as India. It is
possible that during seasonal floods or periods of torrential rainfall, small
lakes situated in low areas of dry land may have spread and connected with
each other, perhaps even connecting down to the sea, which was a
considerable distance to the west of modern Priirtyshja. Under such
circumstances, individual marine animals may have been able to penetrate
beyond their usual habitat through these connected reservoirs or, under
certain conditions, they may have been passively carried to such places.
After the waters receded, their return passage would have been cut off.
Some palaeontologists suggest that floods after heavy rains could have been
so powerful that they engendered turbulent flows that washed slow and
weak animals into lowlands and temporary pools. There could also have
5
Kuznetsov 1958, p. 69-71.
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Preservation for the Musefication of the Natural Heritage Site “Gussinyi Perelet”
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been other natural flows of turbulent water that carried these animals into
the main course of the ancient river.
A special place amongst the Hipparion fauna is reserved for a type of
pinniped (seal) - the semantor - which has only ever been found amongst
the fauna of Gussinyi Perelet. The find again begs the question of how this
animal penetrated such a long way inland. Most likely, it originated where
the delta system flowed into the sea, and it is possible that it migrated in the
same way as the sea turtles.
Some scholars have naturally questioned whether the remains of sea
turtles and semantora might have been redeposited as a result of destructive
Neogene floods and other natural processes, causing the bones of marine
animals to become mixed with terrestrial ones. This, however, is not
supported by the evidence, as it is known that in the process of
redeposition, older sediments tend to undergo significant processing,
becoming rolled-up as a result of displacement, which has not been
observed in this case. Thus, the presence of marine animals such as
pinniped semantora and sea turtles can only be due to the fact that the river
was at some point linked to the sea.
In general, however, the hipparion fauna are typically creatures that
dwelt in lake and river areas in a savannah landscape.
There are thousands of items of skeletal material buried in the
Neogene sand-clay depths of Pavlodar, and most are from ungulates that
may have been victims of a natural disaster. The mass death could have
resulted from a sudden flood caused by rains accompanied by a hurricane; a
wildfire from a lightning strike in the dry summer, which spread across the
steppe; or a long catastrophic drought in which animals died not only from
lack of moisture and food, but attacked each other whilst crowding around
the watering hole. Animals could also have been killed en masse by the
flooding of the ancient river. Skeletal remains of fauna that lived along its
shores were periodically washed away from meadows and rivers banks, and
accumulated in depressions that later formed lens-shaped deposits of bone.
6
The banks of small ponds in the Pavlodar region became burial places
where half-decayed remains of dead animals were periodically covered in
mud. It is interesting to note that about 80% of the hiloteriya rhinoceros
bones discovered there belong to young animals. This is a possible
consequence of temporary droughts which killed off younger animals that
were less able to tolerate lack of water. Flooding could have caused further
burial, but would not have been the cause of death: animals are killed by
floods tend to be buried with their skeletons more-or-less intact, and
6
Orlov 1989, p. 43-44.
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