330
Shamshev & Grootaert: A review of the genus Stilpon from the Oriental region
upper lobe (Fig. 43) undivided, with markedly developed
surstylar comb. Right surstylus (Fig. 44) moderately large,
sublinear, more or less rounded at apex, lacking spines. Left
cercus unbranched, digitiform, fairly long, lacking spines,
with 1 strong ventral bristle in middle part and 1 thinner
marginal bristle basally. Right cercus consisting of two lobes,
lacking spines; left lobe elongate, somewhat broadened
toward apex; right lobe short, tapered toward apex. Phallus
long, hair-like, curved.
Female. Unknown.
Measurements. – Body length 1.4-1.7 mm, wing length 0.9-
1.2 mm.
Etymology. – “Khorng” (curved) “keun” (upwards)
refers to
the fact that the vein r4+5 is strongly curved upward.
Phylogenetic relationships. – The relationships of this species
are unresolved beyond inclusion within the S. divergens
group. Presumably the sister species to S. lekkwar based on
the presence of apical spines on the middle part of the male
left surstylus.
Distribution and seasonal occurrence. – Thailand. Only
known from one locality in Northeast Thailand in the Loei
province. Records are from April till June and one more
record in September.
Stilpon lek, new species
(Figs. 45-49)
Material examined. – Holotype - male,
THAILAND: Loei
province, Na Haeo, near waterfall at FIRS, sample n
°
20013, 23
May.2000, coll. P. Grootaert (SWU).
Paratypes – 1 male, same locality as in holotype, sample n
°
98064,
23.V.1998, coll. P. Grootaert; 2 females, same data as in holotype;
2 males, 1 female, Na Haeo, near stream at FIRS, sample n
°
2026,
27 May.2000, coll. P. Grootaert; 1 male, Na Haeo, near stream on
mud banks at FIRS, sample n
°
21035, 6 May.2001, coll. P. Grootaert;
1 male, 2 females, Na Haeo, FIRS Malaise trap, 18-25 Dec.2001,
coll. P. Grootaert; 1 male, 1 female, same locality as in holotype,
sample n
°
21033, 6 May.2001, coll. P. Grootaert. (SWU, RBINS,
ZRC)
.
Diagnosis. – Species with black thorax, fore tibia yellow.
Male: mid femur with 1 short black anteroventral spine just
beyond middle, abdomen with gland-like structures between
tergites 4-3 and 3-2. Halter contrastingly black.
Description. – Male. Head black in ground-colour, with
minute anterior ocellars, somewhat longer posterior ocellars
and long, cruciate inner verticals. Antenna yellow. Postpedicel
nearly 2.0 times longer than wide. Style about 5 times longer
than postpedicel. Palpus brownish yellow, small,
subtriangular.
Figs. 45-49. Stilpon lek, new species, male. 45, mid leg, anterior view, 46, wing, dorsal view, 47, hypopygium, ventral view, 48, left surstylus,
dorsal view, 49, right surstylus, dorsal view; v br – ventral bristle. Scale bar: 0.1 mm.
331
THE RAFFLES BULLETIN OF ZOOLOGY 2004
Thorax wholly dark brown. Scutum entirely tomentose.
Postpronotal bristle long, black, inclinate. Dorsocentrals in
multiple rows, complete posteriorly. Acrostichals 2-serial,
complete posteriorly.
Legs with colour pattern: hind femur in apical 2/3 yellowish
brown to brownish, fore tarsomere 5 dark brown, mid and
hind tarsomere 5 pale brown; in darker specimens mid and
hind coxae, mid and hind tibiae and mid femur at apex with
brownish tinge. Hind trochanter lacking spines. Mid femur
(Fig. 45) with 4 yellowish ventral bristles in basal half and
1 short black anteroventral spine just beyond middle. Hind
femur (viewed laterally) evenly thickened toward middle;
with row of 5 long anteroventral bristles in apical part and
some prominent dorsal bristles. Fore tibia lacking prominent
bristles. Mid tibia with rows of ventral spinules in apical half
and 1 longer subapical spinule. Hind tibia unmodified.
Wing (Fig. 46) normally developed, covered with uniform
microtrichia; almost uniformly infuscate, somewhat paler
along posterior margin. Costal vein with ordinary short setulae
on anterior margin. Vein R2+3 about 3 times longer than Rs.
Distance between apices of veins R2+3 and R4+5 about 1.3
times longer than distance between apices of veins R1 and
R2+3. R4+5 and M divergent and evenly arcuate in apical
part. Halter with contrast black, elongate knob and pale yellow
stem.
Abdomen largely pale brown, with segments (except segment
8) weakly sclerotized, segment 8 wholly brown. Segments
1-2 unmodified. Narrow gland-like structures present between
tergites 4-3 and 3-2 (Fig. 80). Tergites 3 and 4 shortened.
Almost all tergites with scattered minute brownish setulae,
tergites 7 and 8 with moderately long bristles. Sternites with
similar setation but sternites 3 and 4 bearing 1 pair of inclinate
bristles.
Hypopygium (Fig. 47) dark brown. Hypandrium with 2 strong
bristles in apical part. Epandrium completely divided. Left
epandrial lamella small, fused to hypandrium, bearing 2 short
bristles in apical part. Left surstylus with upper lobe (Fig.
48) undivided, broadened toward apex, with markedly
developed surstylar comb. Right surstylus (Fig. 49) rather
small, rounded apically, sublinear, lacking spines. Left cercus
unbranched, elongate, digitiform, lacking spines, with 1 long
ventral bristle in middle part and several shorter marginal
bristles basally. Right cercus consisting of two lobes, lacking
spines; left lobe elongate, digitiform, somewhat broadened
toward apex; right lobe small, rounded, with 2 long apical
bristles. Phallus long, hair-like, curved.
Female. In most respects identical to male. Fore tarsomere
5 pale brown, mid and hind tarsomere 5 pale yellow. Mid
femur with yellowish ventral bristles only. Mid tibia lacking
spinules. Halter with pale brownish knob. Abdomen lacking
gland-like structures. Terminalia elongate. Segment 8 stronger
sclerotized than preabdomen. Proximal margin of sternite 8
without 2 anteriorly directed rods. Apex of sternite 8 not
partially separated from base. Sternite 10 uniformly
sclerotized, not fused with ventroapical margin of tergite 8.
Cercus elongate oval, brownish yellow, clothed in setulae of
different length.
Measurements. – Body length 1.5-1.9 mm, wing length 0.9-
1.2 mm.
Etymology. – “Lek” means small in Thai and it refers to the
small size of the species
Phylogenetic relationships. – The relationships of this species
are unresolved beyond inclusion within the S. divergens
group.
Distribution and seasonal occurrence. – Thailand. Known
from one locality in Northeast Thailand. Records mainly in
May, but one record also in December.
Stilpon lekkwar, new species
(Figs. 50-53)
Material examined. – Holotype - male,
THAILAND: Loei
province, Na Haeo, near waterfall at FIRS, sample n
°
20013, 23
May.2000, coll. P. Grootaert (coll. RBINS).
Paratypes – 1 male, 17-24 Jun.2001; 2 males, 17-24.IX.99;
1 female, 29 Apr.2001-6 May.2000; 1 female, 8-15 Apr.2001;
1 male, 30 Apr.-7 May.2000; 1 male, 1 female, 7-14
May.2000, same locality as in holotype, coll. P. Grootaert.
2 males, Khring Nam Tok, sample n
°
23041, 20 May.2003,
coll. P. Grootaert; 1 male, Na Haeo FIRS, Malaise trap n
°
2
in bamboo wood, 24 Apr.2003 (leg. P. Grootaert). (SWU,
RBINS, ZRC).
Diagnosis. – Species with black thorax, very similar to
S. lek
but somewhat smaller, hind femur entirely yellow, male
abdomen with gland-like structures between tergites 5-4, 4-
3 and 3-2. Halter pale brown.
Description. – Male. Head black in ground-colour, with
minute anterior ocellars, somewhat longer posterior ocellars
and long, cruciate inner verticals. Antenna yellow. Postpedicel
nearly 2.0 times longer than wide. Style about 5 times longer
than postpedicel. Palpus brownish yellow.
Thorax wholly dark brown. Scutum entirely tomentose.
Postpronotal bristle long, black, inclinate. Dorsocentrals in
multiple rows, complete posteriorly. Acrostichals 2-serial,
complete posteriorly.
Legs with fore tarsomere 5 blackish brown, mid and hind
tarsomere 5 pale brown; otherwise legs yellow. Hind
trochanter lacking spinules. Mid femur (Fig. 50) with row of
4 brownish yellow bristles in basal 1/3, bearing 1 antero- and
1 posteroventral short spines just beyond middle. Hind femur
(viewed laterally) evenly thickened toward middle; with 6-
7 long anteroventral and some prominent dorsal bristles. Fore
tibia lacking prominent bristles. Mid tibia with rows of ventral
spinules in apical half and 1 longer subapical spinule. Hind
tibia unmodified.